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LON-CAPA BRYOPHYTES: Fossombronia
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FOSSOMBRONIA

Raddi - 1818. Jungermanniografia Etrusca. p. 29. (p. 17 of 1841 reprint).
(Type species: Fossombronia angulosa (Dicks.) Raddi)
Each entry of this nomenclator links to the protologue (Gk. protos, first, logos, discourse) which includes everything associated with that name when it was first published. Information from these data are used in the determination of the nomenclatural type for that name, being that element (usually a preserved specimen) to which that name is permanently attached. The abbreviations T:, TL:, D:, B:, and S: refer, respectively, to TYPE specimen, TYPE LOCATIONS (herbarium of deposit), DATA for nomenclatural types, BASIONYM (base name), and STATUS of the taxon, i.e., whether it is accepted, synonomized, or invalid. Invalid names are provided with the appropriate Article number in the International Code of Botanical Nomenclature (Greuter, W. et al. 1994. I.C.B.N. Regnum Vegetabile 131:1 - 389. Entries in [square brackets] include reference to published typifications and corrections to the nomenclator Index Hepaticarum (1962-1990, J. Cramer, Weinheim; Vaduz; Berlin).

Alphabetical Jump Station (by species epithet)

A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

akiensis Horik. - 1934. Botanical Magazine (Shokubutsu-Gaku Zasshi) 48: 453-454.
T: Holotype: Horikawa 11551. February 15, 1933.
TL: HIRO
D: Japan, Honshu, Aki Province, Koi, nr Hiroshima. Margins of cultivated fields and ditches.
S:

alaskana Steere & Inoue - 1974. Bryologist 77: 66-71.
T: Holotype: Steere 72-360. July 20, 1972.
TL: NY; Isotypes: ABSH, TNS
D: U.S.A., Arctic Alaska, Brooks Range, De Long Mountains. On calcareous silt on frost boils. Noluck Lake (68° 47' N Lat., 160° W Long.).
S:

alata G. A. M. Scott & D. C. Pike - 1984. Journal of the Hattori Botanical Laboratory 56: 340-341.
T: Holotype: Pike & Moore s. n. (MUCV 4278). November 2, 1980.
TL: MELU (from MUCV)
D: Australia, Victoria, Lilly Pilly Gully, Wilsons Promontory.
S:

altilamellosa G. A. M. Scott & D. C. Pike - 1987. Journal of the Hattori Botanical Laboratory
62: 367-368.
T: Syntypes: Cargill s. n. (MUCV 4228); Fuhrer, Pike & Scott s. n. (MUCV 6550). September 15, 1990.
TL: MELU (from MUCV)
D: Australia, Victoria and Tasmania.
S: nom. invalid. type non cit. (Art. 37.4)

americana Besch. - Nomen herbarium
T: Specimen: Arsène, November 1906.
TL: NY, PC
D: Mexico, Puebla, l'Aleseca
S: nom. invalid nom. nud (Art. 32.1)

angulosa (Dicks.) Raddi - 1818. Jungermanniografia Etrusca. pp. 29-30. (p.17-18 of 1841 reprint).
T: sub Jungermannia angulosa

B: Jungermannia angulosa Dicks. - 1785. Plantarum Cryptogamicarum Britanniae, Fasciculus 1: 7.
T: Lectotype: Dickson s. n.
TL: BM
D: England
S:

angulosa v. caespitiformis Raddi "caespiformis" - 1818. Jungermanniagrafia Etrusca. p. 30.
(p. 18 of 1841 reprint)
T: Lectotype: Raddi s. n.
TL: PI
D: Italy [the entry "var. caespitiformis Nees - 1838, Naturg. Eur. Leberm." in IH is in error]
S:

angulosa v. dumortieri (Hübener & Genth) Husn. - 1881. Hepaticologia Gallica. Ed. 1. p. 71.
T: sub Codonia dumortieri

areolata G. A. M. Scott & D. C. Pike - 1987. Journal of the Hattori Botanical Laboratory
62: 368-370.
T: Holotype: Pike & Pike, s. n. (MUCV 4628). September 16, 1981.
TL: MELU (from MUCV)
D: Australia, Victoria, Lodge Rd., Grampians.
S:

auricolor G. A. M. Scott & D. C. Pike - 1987. Journal of the Hattori Botanical Laboratory
62: 370.
T: Syntypes: Fuhrer, Pike, & Scott s. n. (MUCV 5435); Fuhrer, Pike &
Scott s. n. (MUCV 5446).
TL: MELU (from MUCV)
D: Australia, South Australia, Wilpena Pound.
S: nom. invalid typ. non cit. (Art. 37.4)

australi-nipponica Horik. - 1934. Journal of Science of Hiroshima University: Series B,
Division 2 (Botany) 2: 138.
T: Holotype: Horikawa 13546. April 10, 1934.
TL: HIRO; Paratype:
D: Formosa (= Taiwan), Taihoku Province, Kelung.
S:

australis Mitt. = Austrofossombronia australis

brasiliensis Steph. - 1900. Species Hepaticarum 1: 382.
T: Lectotype: Ule 109 [Brazil]
TL: G; Paratypes: G- Puiggari 82, 804 [Brazil], Elliott 1258, 1265 [Dominican Republic]
D: Brazil; Dominican Republic [apparently no specimens are extant for the Stephani entry Cuba, Wright]
S: Taxonomic synonym of Fossombronia porphyrorhiza

brasiliensis v. lamellifera R. M. Schust. - 1991. Journal of the Hattori Botanical Laboratory
70: 147.
T: Holotype: Sharp 4077. December 21, 1940
TL: ASBH
D: U.S.A, Oklahoma, Murray County, Prices Falls.
S: nom. invalid hb. non cit. (Art. 37.5)

brasiliensis v. leucorhiza R. M. Schust. - 1991. Journal of the Hattori Botanical Laboratory
70: 147.
T: Holotype: R. M. Schuster 44040. December 19, 1959 (labelled as var. leucotrichia)
TL: F
D: U.S.A., Florida, Torreya State Park.
S: nom. invalid hb. non cit. (Art. 37.5)

caespitiformis De Not. ex Rabenh. - 1860. Hepaticae Europaeae, Die Lebermoose Europa's,
Decas XII und XIV, No. 123, Dresden.
T: Lectotype: Cesati s.n.
TL:
D: Switzerland, Ticino, Locarno.
S:

caespitiformis v. husnotii (Corb.) Corb. - 1904. Revue Bryologique 31: 40.
T: sub Fossombronia husnotii

caespitiformis v. multispira Schiffn. - 1917. Österreichische Botanische Zeitschrift 67: 152.
T: Lectotype: Baumgartner s. n. March 20, 1913.
TL: FH
D: Dalmatien, Gebiet von Zara.
S:

caespitiformis v. subcristata Schiffn. - 1909. Verhandlungen der Kaiserlich-Königlichen
Zoologisch- Botanischen Gesellschaft in Wien 59: 34.
T: Lectotype: Latzel 73
TL: FH
D: Dalmatia, Ragusa, Torrente über dem Dubacpass, 200 m.
S:

caespitiformis v. verrucosa (Lindb.) Müll. Frib. - 1916. Kryptogamen Flora. Ed. 2, 6 (2):
733.
T: sub Fossombronia verrucosa

caledonica Steph. - 1917. Species Hepaticarum 6: 71.
T: Lectotype: Franc s. n. August 17, 1908.
TL: G
D: New Caledonia, Nondoué, Dumbéa, terre humide.
S:

capensis S. W. Arnell - 1952. Botaniska Notiser 1952 : 314.
T: Lectotype: S. W. Arnell, 1876.
TL: S
D: South Africa, Cape Province, Knysna, Bracken Hill Forest.
S:

carinata Gottsche - 1846. Synopsis Hepaticarum. p. 469.
T: Lectotype: Liebmann s. n.
TL: G, Isotype: W
D: Mexico, prope Yavesea, ex hb Gottsche.
S:

cerebriformis G. A. M. Scott & D. C. Pike - 1984. Journal of the Hattori Botanical Laboratory
56: 343.
T: Holotype: Fuhrer, Pike & Scott s. n. (MUCV 4916) August 24, 1982.
TL: MELU (from MUCV)
D: Australia, Victoria, Grampians, Golton's Gorge.
S:

colombica S. Arnell - Nomen herbarium
T: Specimen: Grubb & Guymer B.105. August 4, 1957.
TL: BM
D: Columbia, Sierra Nevada de Cocuy, State of Boyacá. c. 3700 m.
S: nom. invalid nom. nud. (Art. 32.1)

corbulaeformis Trab. in Batt. & Trab. - 1886. Atlas de la Flore d'Alger. Fasc. 1: 7.
T: Lectotype:
TL:
D: Algeria.
S:

crassifolia Spruce - 1885. Transactions and Proceedings of the Botanical Society of Edinburgh
15: 527.
T: Lectotype: Spruce s. n.
TL: G
D: Ecuador, Baños, ad pedem M. Tunguragua.
S:

crispa Nees - 1846. Synopsis Hepaticarum. p. 469.
T: Lectotype: ?Zeyher [vide Perold, Bothalia 27: 137. 1997.]
TL: STR
D: South Africa, Cape of Good Hope.
S:

cristata Lindb. - 1874. Notiser ur Sällskapets pro Fauna et Flora Fennica Förhandlingar 13: 388.
T: Lectotype: Lindberg s. n. September 29, 1873
TL: H
D: Finland, Helsinki.

cristata v. loitlesbergeri (Schiffn.) Chalaud - 1930. Revue Générale de Botanique 42: 569.
T: sub Fossombronia loitlesbergeri.

cristata v. wondraczekii (Corda) Lindb. - 1874. Notiser ur Sällskapets pro Fauna et Flora
Fennica Förhandlingar 13: 389.
T: sub Jungermannia wondraczekii

cristula Austin - 1869. Proceedings of the Academy of Natural Sciences of Philadelphia (1869)
21: 228.
T: Lectotype: Austin, Hep. Bor.- Amer. no. 121. October, 1868.
TL: MANCH
D: U.S.A., New Jersey, nr Batsto.
S:

cristula v. verdoornii Chalaud - 1931. Annales Bryologici 4: 143.
T: Lectotype: Verdoorn s. n. Hep. Sel. Crit. 59. April, 1930.
TL:
D: Java, Tjibodas, in horto botanic.
S:

crozalsii Corb. - 1903. Revue Bryologique 30: 13.
T: Lectotype: Crozals s. n. April 27, 1902.
TL:
D: France, Hérault, Commune de Lieuran-les-Béziers, bois du château de Ribaute.
S:

cubana Gottsche ex Austin - 1876. Botanical Bulletin 1: 36. [The title Botanical Gazette in IH is in error]
T: Lectotype: Wright
TL: G
D: Cuba
S:

cultriformis G. A. M. Scott & D. C. Pike - 1987. Journal of the Hattori Botanical Laboratory
62: 371.
T: Holotype: Pike & Fuhrer s. n. (MUCV 6310). September 8, 1985.
TL: MELU (from MUCV); Paratypes: MUCV 6302, 6303, 6301, 6307, 6308, 6309, 6311
D: Australia, South Australia, The Coorong, 1 km from Cooke's plain turn-off nr Meningee.
S:

densa G. A. M. Scott & D. C. Pike - 1987. Journal of the Hattori Botanical Laboratory 62: 372.
T: Holotype: Fuhrer s. n. (MUCV 2455). September 5, 1974
TL: MELU (from MUCV)
D: Australia, Victoria, State Forest, Rushworth.
S:

densilamellata S. W. Arnell - 1952. Botaniska Notiser (1952): 317.
T: Lectotype:
TL: S
D: South Africa, Cape Province, Lions Head, nr Kloofnek.
S:

dentata Steph. - 1900. Species Hepaticarum 1: 394.
T: Lectotype: Weymouth s. n. Sept. 22, 1893.
TL: G
D: Australia, Tasmania, Trevallyn, Launceston, nr bridge, on earth.
S:

dentata f. robusta Rodway - 1918. Papers and Proceedings of the Royal Society of Tasmania
(1917): 106.
T: Lectotype: Rodway
TL:
D: Australia, Tasmania, Cradle Mountain.
S:

dumortieri (Hübener & Genth) Lindb. - 1874. Notiser ur Sällskapets pro Fauna et Flora Fennica
Förhandlingar 13: 417.
T: sub Codonia dumortieri.

B: Codonia dumortieri Hübener & Genth - 1837. Deutschlands Lebermoose in
Getrockneten Exemplare Fasc. 4, No. 80.
T: Lectotype: Lindberg. September, 1863. [vide Grolle,]
TL: H-SOL
D: Sweden, Uppland, Lindigo (nr Stockholm).
S: nom. invalid nom. nud. (Art. 32.1) = Fossombronia foveolata

echinata Macvicar - 1911. Revue Bryologique 38: 73.
T: Lectotype: Nicholson. April, 1910.
TL: CBG
D: Italy, Amalfi.
S:

erythrorhiza Hampe - Nomen herbarium
T: Specimen:
TL:
D:
S: nom. invalid nom. nud. (Art. 32.1)

fernandeziensis Steph. - 1911. Kongliga Svenska Vetenskapaakademiens Handligar 46: 15.
T: Lectotype: Skottsberg 104. 1908
TL: G
D: Juan Fernandez, Masatierra.
S:

fimbriata Paton - 1974. Journal of Bryology 8: 1.
T: Holotype: Paton s. n. August, 1968.
TL: BM
D: Scotland, Mid- Perth, Glen Lochay, On moist schistose soil beside the hydro road above Badour.
S:

fleischeri Osterwald ex Loeske - 1928. Verhandlungen des Botanischen Vereins der Provinz
Brandenburg 70: 125. [pp. 1-3 of reprint.]
T: Lectotype: Osterwald s. n. September, 1910.
TL:
D: Germany, Ausstich bei Buch, bei Berlin, 60 m.
S:

foreaui Udar & S. C. Srivast. - 1973. Beihefte zur Nova Hedwigia 47: 463.
T: Holotype: Udar & S. C. Srivastava 38F/66. January 4, 1966.
TL: LWU
D: India, Lake side road Kodaikanal, ca. 6,854 ft.
S:

forsythii Steph. 1917. Species Hepaticarum 6: 72.
T: Lectotype: Forsythe 51. September 23, 1898.
TL: G
D: Australia, Nepean river.
S:

foveolata Lindb. - 1873. Helsingfors Dagblad, Dec. 28. p. 353. [The entry Notiser
ur Sällskapets pro Fauna et Flora Fennica Förhandlingar 13: 382 in IH is in error.]
T: Lectotype: Lindberg s. n. September, 1863 [vide Grolle, Trans. Brit. Bryol. Soc. 5: 767. 1969.]
TL: H-SOL
D: Sweden, Uppland, ins. Lindigo [nr Stockholm].
S:

foveolata v. cristula (Austin) R. M. Schust. - 1992. The Hepaticae and Anthocerotae of North
America East of the Hundredth Meridian. Volume 5. p. 383.
T: sub Fossombronia cristula

fuhreri G. A. M. Scott & D. C. Pike - 1987. Journal of the Hattori Botanical Laboratory 62: 374.
T: Holotype: Pike & Fuhrer s. n., MUCV 5729. September 10, 1984.
TL: MELU (from MUCV); Paratypes: MUCV 5723, 5724, 5730, 5731.
D: Australia, Victoria, Portland, Point Danger. In crevices of volcanic rock on cliff near sea.
S:

gigantea Steph. - 1900. Species Hepaticarum 1: 391.
T: Lectotype: Knight
TL: G; Paratype: Kuk 461 p.p. (G)
D: New Zealand, ex Herb. Gottsche; Paratype: New Zealand, Nelson.
S:

grandis Steph. - 1900. Species Hepaticarum 1: 383.
T: Lectotype: Volkens 1226 p.p. October 9, 1893.
TL: G
D: Kilimandscharo.
S:

gregaria Colenso - 1888. Transactions and Proceedings of the New Zealand Institute 20: 252.
T: Lectotype: Hep., a 1375.
TL: BM
D: New Zealand, woods at Tarawera, hilly country north of Napier,
1886: Mr. H. Hill. [These data are taken from the protologue, the
specimen in BM is labelled Hep., a 1375 Col.]
S:

grossepapillata Steph. - 1914. Journal and Proceedings of the Royal Society of New South
Wales 48: 105.
T: Lectotype: Watts 988. September - October, 1906.
TL: G
D: Australia, New South Wales, edge of mining hole, nr Young.
S:

hamato-hirta Steph. - 1894. Hedwigia 33: 8.
T: Lectotype: Gordon 108. August, 1889.
TL: G
D: Ascension Island, Green Mountain, 2000 ft.
S:

herzogii K. I. Goebel in Herzog - 1916. Bibliotheca Botanica 87: 269.
T: Lectotype: Herzog 4388. 1911.
TL: JENA
D: Bolivia, Cerros de Malaga.
S:

hewsonii G. A. M. Scott & D. C. Pike - 1987. Journal of the Hattori Botanical Laboratory
62: 375.
T: Holotype: Hewson s. n., MUCV 5492. August 22, 1984.
TL: MELU (from MUCV)
D: Australia, Western Australia, W. of Moora, Dandaragan Road.
S:

himalayensis Kashyap - 1915. New Phytologist 14: 4.
T: Lectotype: Kashyap
TL:
D: India, Mussourie.
S:

histrix - Nomen herbarium
T: Specimen:
TL:
D:
S: nom. invalid nom. nud. (Art. 32.1).

hispidissima Steph. - 1900. Species Hepaticarum 1: 389.
T: Lectotype: Bolander 59.
TL: G
D: U.S.A., California, San Francisco.
S:

husnotii Corb. - 1889 Mémoires de la Socié Nationale des Sciences Naturelles et Mathématiques
de Cherbourg 26: 353 [The entry 1890. Revue Bryologique 17: 3 in IH is in error.]
T: Lectotype: Corbière
TL:
D: France, Maupertus près Cherbourg.
S:

husnotii v. anglica Nicholson - 1914. Journal of Botany, British and Foreign 52: 106.
T: Lectotype: Nicholson s. n. March 23, 1913.
TL: CBG
D: England, S. Devon, Babbacombe Bay, moist ground nr sea.
S:

incurva Lindb. - 1873 Helsingfors Dagblad (273) pp. 2-3: October 7.
T: Lectotype: Lindberg s. n. Sepetember, 1873 [vide Grolle, Feddes Repertorium 87: 192. 1976.]
TL: H-SOL
D: Finland, Fredriksburg, nr Helsingfors.
S:

incurva v. tenera Lindb. - 1874. Notiser ur Sällskapets pro Fauna et Flora Fennica
Förhandlingar 13: 382.
T: Lectotype: Lindberg s. n. September, 1873
TL: H-SOL
D: Finland, Fredriksburg, nr Helsingfors.
S:

indica Steph. - 1917. Species Hepaticarum 6: 73.
T: Lectotype: Pfleiderer s. n. August 1906.
TL: G
D: India, South Canara, Mangalore, hb Levier 5010.
S:

integerrima Steph. - 1900. Species Hepaticarum 1: 394.
T: Lectotype: Colenso 1376 p.p.
TL: G
D: New Zealand.
S:

integrifolia Steph. - 1917. Species Hepaticarum 6: 73.
T: Lectotype: Cheel s. n. August 5, 1900.
TL: G
D: Australia, New South Wales, Rockwood, nr Sydney, ex Herb. Watts.
S:

intermedia Corb. ex Trab. - 1942. Revue Bryologique et Lichénologique 12: 25.
T: Lectotype:
TL:
D: Algeria
S:

intestinalis Taylor - 1846. London Journal of Botany 5: 408.
T: Lectotype: Drummond s. n.
TL:
D: Australia, Swan River.
S:

japonica Schiffn. - 1899. Österreichische Botanische Zeitschrift 49: 389.
T: Lectotype: Miyake 50. November, 1897.
TL: FH; Isotypes: TNS, W
D: Japan, Tokyo, Botanical Garden.
S:

kashyapii S. C. Srivast. & Udar - 1975. Nova Hedwigia 26: 816.
T: Holotype: Srivastava 10F/67. October, 1967.
TL: LWU
D: India, Western Himalayas, Rahla, ca. 9,000 ft, on way to Rohtang pass in Kulu valley, about 12 miles from Manali.
S:

laciniata G. A. M. Scott & D. C. Pike - 1987. Journal of the Hattori Botanical Laboratory
62: 375.
T: Holotype: Fuhrer & Scott s. n., MUCV 5402. August 30, 1984.
TL: MELU (from MUCV)
D: Australia, 1 km east of of Salt Creek, The Coorong.
S:

lamellata Steph. - 1894. Hedwigia 33: 9.
T: Lectotype: Hauthal s. n.
TL:
D: Argentina, Buenos Aires, ex hb. O. Kuntze.
S:

latifolia Steph. - Nomen herbariorum
T: Specimen: Pfleiderer 32
TL: G
D: India, Orientalis.
S: nom. invalid nom. nud. (Art. 32.1).

leucoxantha (Lehm.) Lehm. & Lindenb. - 1846. Synopsis Hepaticarum. p. 469.
T: sub Jungermannia leucoxantha

B: Jungermannia leucoxantha Lehm. - 1829. Linnaea 4: 368
T: Lectotype: Ecklon s. n.
TL: S; Isotype: BM
D: South Africa, Cape of Good Hope, Table Mountain.
S:

levieri Steph. - 1917. Species Hepaticarum 6: 74.
T: Lectotype: Gollan s. n.
TL: G
D: India, Himalayas, Mussoorie, 5500 ft.
S:

loitlesbergeri Schiffn. - 1909. Hedwigia 48: 195.
T: Lectotype: Baumgartner & Loitlesberger s. n. April 1906 & March 1907. V Schiffner, Hepaticae europaeae exsiccatae 1340.
TL: FH
D: Dalmatia. Insel Arbe (Rab), in Walde capo Fronte.
S:

longiseta (Austin) Austin - 1873. Hepaticae Boreali-Americanae 118. p. 30.
T: sub Androcryphia longiseta

B: Androcryphia longiseta Austin - 1869. Proceedings of the Academy of Natural Sciences of Philadelphia 21: 228.
T: Lectotype: Bolander
TL: MANCH
D: U.S.A, California [Texas, Wright in IH is in error = F. texana]
S:

lophoclada Spruce - 1885. Transactions and Proceedings of the Botanical Society of Edinburgh
15: 529.
T: Lecotype: Spruce s. n.
TL: G
D: Peru, Andes peruviana, Monte Campana, supra terra purpurea.
S:

luetzelburgiana K. I. Goebel - 1912. Flora 105: 55.
T: Lectotype:
TL:
D:
S:

macounii Austin - 1876. Botanical Bulletin 1: 36. [The title Botanical Gazette in IH is in error]
T: Lectotype: Macoun s. n. September, 1875.
TL: NY
D: Canada, Portage, La Lochs, Lat. 57.
S:

macrocalyx Steph. - 1917. Species Hepaticarum 6: 74.
T: Lectotype: Franc s. n. June 28, 1909.
TL: G
D: New Caledonia, Pont du Diable près Tonghoué, 150 m.
S:

macrophylla Colenso - 1886. Transactions and Proceedings of the New Zealand Institute
18: 285.
T: Lectotype: Hep., a 1374. Col.
TL: BM
D: New Zealand, County of Waipawa, east slopes of Ruahine Mountain Range, damp shaded
spots, ravines. [These data are taken from the protologue, the specimen in BM is labelled only
"Hep., a 1374 Col."]
S:

magnaspora G. A. M. Scott & D. C. Pike - 1987. Journal of the Hattori Botanical Laboratory
62: 377.
T: Syntypes: Fuhrer, Pike & Scott s. n. MUCV 7320. June 20, 1984; Pike &
Pike s. n., MUCV 4626. October 16, 1981.
TL: MELU (from MUCV)
D: Australia, Victoria, Grampians, Glenely Forest.
S: nom. invalid type non cit. (Art. 37.4)

maritima G. A. M. Scott & D. C. Pike - 1987. Journal of the Hattori Botanical Laboratory
62: 378.
T: Syntypes: Fuhrer & Pike s. n., MUCV 5518, 5519. September 15, 1984.
TL: MELU (from MUCV)
D: Australia, Victoria, Portland, Nelson Bay.
S: nom. invalid type non cit. (Art. 37.4)

maritima (Paton) Paton - 1994. Journal of Bryology 18: 367.
T: sub Fossombronia pusilla v. maritima

mauritanica Trab. - 1942. Revue Bryologique et Lichénologique 12: 25.
T: Lectotype: Castiglione s. n.
TL:
D: Algeria.
S:

mexicana Steph. - 1917. Species Hepaticarum 6: 74.
T: Lectotype: Arsèn 7755.
TL: G, Paratypes: G.
D: Mexico, Puebla [Arsène], ex Herb. Lacoutre. Two packets in G labelled Fossombronia mexicana St. ms., Mexico, Arsèn[e] are designated as paratypes, the designated lectotype packet is labelled "Fossombronia mexicana St. nsp."
S:

microlamellata G. A. M. Scott & D. C. Pike 1984. Journal of the Hattori Botanical Laboratory
56: 347.
T: Holotype: Pike & Fuhrer s. n. MUCV 4926. September 13, 1982.
TL: MELU (from MUCV); Isotype: G
D: Australia, Victoria, South Belgrave, Courtney's Road, on bare soil on a suburban Nature
Strip.
S:

microspora Herzog - 1935. Transactions and Proceedings of the Royal Society of New Zealand
65: 356.
T: Lecotype: Allison H 346. December 21, 1930.
TL: JENA
D: New Zealand, North Island, nr Atiamuri, 1000 ft., stream bank, in shade.
S:

mittenii Tind. - 1898. Journal of Botany, British and Foreign 36: 44.
T: Lectotype: Mitten. August, 1875.
TL: NY
D: England, North Devon, bank on side of the road between Parracombe and Braunton.
S:

mittenii v. echinata (Macvicar) Macvicar ex Müll. Frib. - 1954. Rabenhorst's Kryptogamen Flora.
Ed. 3, 6: 542.
T: sub Fossombronia echinata

montaguensis S. W. Arnell - 1952. Botaniska Notiser (1952): 316.
T: Lectotype:
TL: S
D: South Africa, Cape Province, Montagu.
S:

mylioides Inoue - 1973. Journal of the Hattori Botanical Laboratory 37: 295-297.
T: Holotype: Inoue 21055. January 14, 1973.
TL: TNS
D: Japan, Ryukyu Isls., Iriomote Isl., along the River Urauchi: nr Gampire Falls, on moist rocks.
S:

naumannii Schiffn. - 1889. Die Forschungsreise S. M. S. "Gazelle", 4, Botanik,
No. 4. p. 39.
T: Lectotype: Naumann s. n. December, 1874.
TL: FH; Isotype: G
D: Kerguelens (France), Successful Bay (Gazelle Expedition).
S:

naumannii v. rielloides Schiffn. - 1889. Die Forschungsreise S. M. S. "Gazelle", 4, Botanik,
No. 4. p. 40.
T: Lectotype: Naumann s. n. December, 1874.
TL: FH; Isotype: G
D: Kerguelens (France), Successful Bay (Gazelle Expedition).
S:

nigricaulis Colenso - 1886. Transactions and Proceedings of the New Zealand Institute 18: 248.
T: Lectotype: Hep., a 1377 Col.
TL: BM
D: New Zealand, Norsewood, Coutny of Waipawa, nr the Mangatawhainni River,
cliffy sides of road. [These data are taken from the protologue, the specimen at BM is
labelled only "Hep., a 1377 Col."]
S:

occidento-africana S. W. Arnell - 1952. Botaniska Notiser (1952): 317.
T: Lectotype:
TL: S
D: Sierra Leone.
S:

papillata Steph. - 1889. Hedwigia 28: 157.
T: Lectotype: Wild s. n.
TL: G
D: Australia, Queensland, ex Herb. Brotherus 19.
S:

paranapanemae Schiffn. ex. S. W. Arnell - 1964. Österreichische Akademie der Wissenschaften
Mathematisch - Naturwissenschaftliche Klasse, Denkschriften 111: 34.
T: Lectotype:
TL:
D: Brazil, São Paulo.
S:

perpusilla (Colenso) Steph. - 1900. Species Hepaticarum 1: 379.
T: sub Noteroclada perpusilla

B: Noteroclada perpusilla Colenso - 1885, Transactions and Proceedings of the New Zealand Institute 17: 260.
T: Lectotype: Colenso 2019.
TL: G
D: New Zealand.
S:

peruviana Gottsche & Hampe = Austrofossombronia peruviana.

porphyrorhiza (Nees) Prosk. - 1955. Bryologist 58: 197.
T: sub Jungermannia porphyrorhiza
B: Jungermannia porphyrorhiza Nees in Mart.
T: Lectotype: Martius s. n.
TL: STR
D: Brazil.
S:

ptychophylla Spruce - 1885. Transactions and Proceedings of the Botanical Society of
Edinburgh 15: 529.
T: Lectotype: Spruce s. n. January, 1859.
TL: MANCH
D: Ecuador, Mt. Altar, Condorasto, 3700 m.
S: Taxonomic synonym of Austrofossombronia peruviana

pulvinata Steph. - 1911. Wissenschaftliche Ergebnisse der Deutschen Zentral-Afrika Expedition,
1907-1908, II (Botanik). p. 113.
T: Lectotype: Mildbraed 810.
TL: G
D: Ruanda [Rwanda], Kiwu See [Kivo Lake].
S:

punctata G. A. M. Scott & D. C. Pike - 1984. Journal of the Hattori Botanical Laboratory
56: 341.
T: Holotype: Scott, MUCV 2485. August, 1974.
TL: MELU (from MUCV)
D: Australia, Victoria, Wyperfeld National Park.
S:

purpureospora G. A. M. Scott & D. C. Pike - 1987. Journal of the Hattori Botanical Laboratory
62: 379.
T: Syntypes: Fuhrer, Pike & Scott s. n. MUCV 5431. August 26, 1986;
Fuhrer & Pike s. n. MUCV 6281. August 31, 1985.
TL: MELU (from MUCV)
D: Australia, Victoria.
S: nom. invalid type non cit. (Art. 37.4)

pusilla (L.) Nees - 1838. Naturgeschichte der Europäischen Lebermoose. Vol. 3. p. 319.
T: sub Jungermannia pusilla L.

B: Jungermannia pusilla L. - 1753. Species Plantarum Vol. 2. p. 1136
T: Lectotype: Dillenius, Historia Muscorum, t.74, f.46.
TL: LINN; Epitype OXF
D: ?England
S:

pusilla var. capitata Nees - 1838. Naturgeschichte der Europäischen Lebermoose. Vol. 3. p. 320.
T: Lectotype: Coll. Ignota
TL: STR
D: In fossis agrorum sylvatio.
S:

pusilla ssp. cristatella R. M. Schust. - Nomen herbariorum.
T: Specimen: Schuster M-438. April 10, 1951.
TL: G
D: U.S.A., Mississippi, Lafayette Co., 15-16 miles S of Oxford.
S: nom. invalid nom. nud. (Art. 32.1)

pusilla v. corsa Lindb. - Nomen Herbariorum
T: Specimen: Hjelt s. n. Feb 2, 1876.
TL: G
D: Corsica, Ajaccio, arena humida in montibus.
S: nom. invalid nom. nud. (Art. 32.1)

pusilla v. decipiens Corb. - 1889. Mémoires de la Société Nationale des Sciences Naturelles et
Mathématiques de Cherbourg 26: 355.
T: Lectotype: Corbière s. n.
TL:
D: France, Octeville, environs de la Fauconnière or Equeurdreville, près du hameau Guerry.
S:

pusilla v. ochrospora Lindb. - 1874. Notiser ur Sällskapets pro Fauna et Flora Fennica
Förhandlingar 13: 387.
T: Lectotype: Lindberg s. n. May 1, 1869.
TL: H-SOL
D: Great Britain, Lewis Park.
S:

pusilla v. maritima Paton - 1973. Journal of Bryology 7: 244.
T: Holotype: Paton s. n. March, 1969.
TL: BM
D: British Isles, Channel Islands, Jersey, St. Brelade, W of Noirmont Point, peaty soil on cliff top.
S: Nomenclatural synonym of Fossombronia maritima.

reticulata Steph. - 1894. Hedwigia 33: 9.
T: Lectotype: Kirk 62
TL: G
D: New Zealand, nr Auckland.
S:

rosulata Colenso - 1886. Transactions and Proceedings of the New Zealand Institute 18: 248.
T: Lectotype: Hep., a 1376. Col.
TL: BM
D: New Zealand, River Mangatawhainui, County of Waipawa, Seventy Mile Bush. [These data are taken from the protologue, the specimen in BM is labelled only "Hep., a 1376. Col."]
S:

rudis G. A. M. Scott & D. C. Pike - 1988. Beiheft zur Nova Hedwigia 90: 110.
T: Holotype: Fuhrer, Pike & Pike s. n. MUCV 6435. July 25, 1985.
TL: MELU (from MUCV)
D: Australia, Northern Territory, Stoke's Creek, nr King's Canyon, description
from cultivated specimen MUCV 7313.
S:

rupestris G. A. M. Scott & D. C. Pike - 1987. Journal of the Hattori Botanical Laboratory
62: 381.
T: Syntypes: Fuhrer, Pike & Scott s. n. MUCV 5447. Plants cultivated as MUCV 7335. August 26, 1984.
TL: MELU (from MUCV)
D: Australia, Wilpena Pound.
S: nom. invalid type non cit. (Art. 37.4)

salina Lindb. - 1875. Acta Societatis Scientiarum Fennicae 10: 533.
T: Lectotype: Austin, N. American Hepaticae No. 119.
TL:
D: U.S.A.
S: nom. invalid nom. nud. (Art. 32.1)

scrobiculata G. A. M. Scott & D. C. Pike - 1984. Journal of the Hattori Botanical Laboratory
56: 341.
T: Holotype: Fuhrer s. n. MUCV 4270. July 25, 1973
TL: MELU (from MUCV)
D: Australia, Victoria, Rushworth.
S:

spinifolia Steph. - 1900. Species Hepaticarum 1: 389.
T: Lectotype: Breutel s. n.
TL: G
D: South Africa, Gnadenthal, Halensis, ex hb K. Müller.
S:

stephanii Schiffn. ex Steph. - 1900. Species Hepaticarum 1: 381.
T: Lectotype: de l'Isle 320. 1875.
TL: G
D: Mascarene Islands, Ile de la Réunion (France).
S:

subsaccata Steph. - 1917. Species Hepaticarum 6: 75.
T: Lectotype: Lesquereux s. n.
TL: G
D: Australia, Lookout Mountains, ex hb Pearson. 1850.
S:

tenuifolia Spruce - 1885. Transactions and Proceedings of the Botanical Society of Edinburgh
15: 528.
T: Lectotype: Spruce s. n.
TL: G
D: Ecuador, Andes Quitenses, in Mte. Altar.
S:

tesselata G. A. M. Scott & D. C. Pike - 1987. Journal of the Hattori Botanical Laboratory
62: 382.
T: Holotype: Fuhrer & Pike s. n. MUCV 6402. August 31, 1985.
TL: MELU (from MUCV)
D: Australia, Grampians, Victoria Valley Road, nr Serra Road junction, on roadside.
S:

texana Lindb. - 1875. Acta Societatis Scientiarum Fennicae 10: 533.
T: Lectotype: Wright s. n.
TL: H-SOL
D: U.S.A., Texas, Wright [The entry California, Bolander in IH is in error = F. longiseta]
S:

trinitensis Mitten - Nomen herbarium
T: Specimen:
TL: NY
D:
S: nom. invalid nom. nud. (Art. 32.1)

truncata G. A. M. Scott & D. C. Pike - 1987. Journal of the Hattori Botanical Laboratory
62: 383.
T: Syntypes: Pike & Scott s. n. MUCV 5180. June 20, 1984; Fuhrer &
Pike s. n. MUCV 4918. September 16, 1982
TL: MELU (from MUCV)
D: Australia, Victoria.
S: nom. invalid type non cit. (Art. 37.4)

tuberifera K. I. Goebel - 1898. Organographie der Pflanzen. p. 292.
T: Lectotype: Goebel
TL:
D: Venezuela, Tovar.
S:

tumida Mitt. - 1877 "1888.". Journal of the Linnean Society. Botany 16: 193.
T: Lectotype: Eaton s. n.
TL: BM
D: South Africa, Cape Town.
S:

tylimanthoidea Steph. Nomen herbariorum.
T: Specimen: Keys 150.
TL: G
D: Australia, ex Herb. Melbourne.
S: nom. invalid nom. nud. (Art. 32.1)

vermiculata G. A. M. Scott & D. C. Pike - 1984. Journal of the Hattori Botanical Laboratory
56: 345.
T: Holotype: Scott s. n. MUCV 676. July 28, 1971.
TL: MELU (from MUCV)
D: Australia, Victoria, W. of Bendigo, nr Bealiba.
S:

verrucosa Lindb. - 1874. Notiser ur Sällskapets pro Fauna et Flora Fennica Förhandlingar
13: 386.
T: Lectotype: Paris s. n. Gottsche & Rabenhorst Hep. Eur. 439. I, 1867.
TL: H-SOL
D: Algeria, Belida, Mouzaiaville.
S:

wattsii Steph. - 1917. Species Hepaticarum 6: 75.
T: Lectotype: Watts 881. September, 1905.
TL:
D: Australia, New South Wales, The Gap nr Young.
S:

wondraczekii (Corda) Dumort. - 1835. Recueil d'Obsrvations sur les Jungermanniacées.
Fascicle I. p. 11.
T: sub Jungermannia wondraczekii

B: Jungermannia wondraczekii Corda in Sturm - 1830. Deutschlands Flora,
Abteilung II, Cryptogamie, Heft 19-20 [= Deutschlands Jungermannien,
Heft 1-2]: 30.
T: Lectotype: Wondraczek s.n.
TL: PR
D: Czech Republic, Tiergarten Stern bei Prag.
S:

wondraczekii v. loitlesbergeri (Schiffn.) Chalaud ex Müll. Frib. - 1954. Kryptogamen Flora
Ed. 3, 6: 545.
T: sub Fossombronia loitlesbergeri

wrightii Austin - 1876. Botanical Bulletin 1: 36. [The listing Botanical Gazette in IH is in error]
T: Lectotype: Wright
TL:
D: Cuba.